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Crocodiles (family Crocodylidae) or true crocodiles are large, semiaquatic that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include all extant taxon members of the order Crocodilia, which includes the and (both members of the family Alligatoridae), the gharial and false gharial (both members of the family Gavialidae) as well as other extinct Taxon.
Crocodile Measurement, morphology, behaviour and ecology differ among species. However, they have many similarities in these areas as well. All crocodiles are semiaquatic and tend to congregate in freshwater habitats such as , , and sometimes in brackish water and Seawater. They are carnivorous animals, feeding mostly on such as fish, , and , and sometimes on such as and , depending on species and age. All crocodiles are tropical species that, unlike alligators, are very sensitive to cold. Many species are at the risk of extinction, some being classified as critically endangered.
A very early extant Ancient Greek source is an Aesop's Fable named Ἀλώπηξ καὶ κροκόδειλος of the sixth century BC. Herodotus a century after in consideration of the Greek word thought it was from the Ionic Greek period.
The Latin language word crocodilus existed in first century AD in the work Naturalis Historia of Pliny the Elder. Examples in writing in the sixth and seventh century AD, crocodillorum and crocodili existed, though corcodrillus and cocodrillus were forms in Medieval Latin. The Latin form is found as cocodrille c. the 13th century in the Old French work Li livres dou tresor, in which the croc. is jaune (yellow).
The earliest known source in the English language is within the work Kyng Alisaunder King Alisaunder Weber edition (1810) from towards the beginning of the 14th century, a magical romance poetry work of rhyming couplets, where the word is found line 5720:
Writing sometime after 1483 on his visit to the Felix Fabri in German and Latin mentions the Cocodrillen and cocodrillos respectively. By 1538 the exact same form of the modern word as the current English is found in French; croc. is within a poem of Edmund Spenser, thought written in 1579–80, and in the works of William Shakespeare, who died in 1616. The similitude of the English word formation to a Latin source was caused at least by the transmission of a relevant ancient science from the influx of the publication of translations of Pliny the Elder some time towards the end of fourteenth and, or, beginning of fifteenth century. The publication of concrete realization in the anatomical work of Andreas Vesalius during 1543 inspired the creation of monographs and books of animals contributing to new science in zoology.
Although crocodiles, alligators, and the gharial are similar in appearance, they belong to separate biological families. The gharial, with its narrow snout, is easier to distinguish, while morphological differences are more difficult to spot in crocodiles and alligators. The most obvious external differences are visible in the head, with crocodiles having narrower and longer heads, with a more V-shaped than a U-shaped snout compared to alligators and caimans. Another obvious trait is that the upper and lower jaws of the crocodiles are the same width, and the teeth in the lower jaw fall along the edge or outside the upper jaw when the mouth is closed; therefore, all teeth are visible, unlike an alligator, which possesses in the upper jaw small depressions into which the lower teeth fit. Also, when the crocodile's mouth is closed, the large fourth tooth in the lower jaw fits into a constriction in the upper jaw. For hard-to-distinguish specimens, the protruding tooth is the most reliable feature to define the species' family. Crocodiles have more webbing on the toes of the hind Foot and can better tolerate saline water due to specialized for filtering out salt, which are present, but non-functioning, in alligators. Another trait that separates crocodiles from other crocodilians is their much higher levels of aggression.
Crocodylidae is defined as a crown group composed of the last common ancestor of the Nile crocodile ( Crocodylus niloticus), the Dwarf crocodile ( Osteolaemus tetraspis), and all of its descendants. It contains two subfamilies: Crocodylinae and Osteolaeminae. Crocodylinae contains 13–14 living species, as well as 6 extinct species. Osteolaeminae was named by Christopher Brochu in 2003 as a subfamily of Crocodylidae separate from Crocodylinae, and contains the two extant taxon genus Osteolaemus and Mecistops, along with several extinct genera. The number of extant species within Osteolaeminae is currently in question.
Below is a cladogram showing the relationships of the major extant taxon crocodile groups based on molecular studies, excluding separate extinct taxa:
Below is a more detailed cladogram of Crocodylidae, based on a 2021 study using paleogenomics that extracted DNA from the extinct Voay. Recently recognised species ( M. leptorhynchus, C. halli and the third Osteolaemus species) placed according to 2023 study by Sales-Oliveira et al.
Alternatively, some morphological studies have recovered Mecistops as a basal member of Crocodylinae, more closely related to Crocodylus than to Osteolaemus and the other members of Osteolaeminae, as shown in the cladogram below.
Like other , crocodilians are diapsid, although their post-temporal fenestrae are reduced. The walls of the braincase are bony but lack supratemporal and postfrontal bones.Grigg, Gordon and Gans, Carl (1993) Morphology And Physiology Of The Crocodylia , in Fauna of Australia Vol 2A Amphibia and Reptilia, chapter 40, pp. 326–336. Australian Government Publishing Service, Canberra. Their are not free, but held in place by a membrane that limits movement; as a result, crocodiles are unable to stick out their tongues. Crocodiles have smooth skin on their bellies and sides, while their dorsal surfaces are armoured with large osteoderms. The armoured skin has scales and is thick and rugged, providing some protection. They are still able to absorb heat through this armour, as a network of small capillaries allows blood through the scales to absorb heat. The osteoderms are highly vascularised and aid in calcium balance, both to neutralize acids while the animal cannot breathe underwater and to provide calcium for eggshell formation. Crocodilian tegument have pores believed to be sensory in function, analogous to the lateral line in fishes. They are particularly seen on their upper and lower jaws. Another possibility is that they are secretory, as they produce an oily substance which appears to flush mud off.
The brain volume of two adult crocodiles was 5.6 cm3 for a spectacled caiman and 8.5 cm3 for a larger Nile crocodile.
The largest crocodile ever held in captivity is a saltwater–Siamese hybrid named Yai (, meaning big; 10 June 1972 – 15 August 2025) at the Samutprakarn Crocodile Farm and Zoo, Thailand. This animal measures in length and weighs . The Guinness Book of Records. Largest Captive Crocodile , Worldcrocodile.com
The longest crocodile captured alive was Lolong, a saltwater crocodile which was measured at and weighed at by a National Geographic team in Agusan del Sur Province, Philippines.
Chemoreception in crocodiles is especially interesting because they hunt in both terrestrial and aquatic surroundings. Crocodiles have only one olfactory chamber and the vomeronasal organ is absent in the adults indicating all olfactory perception is limited to the olfactory system. Behavioural and olfactometer experiments indicate that crocodiles detect both air-borne and water-soluble chemicals and use their olfactory system for hunting. When above water, crocodiles enhance their ability to detect volatile odorants by gular pumping, a rhythmic movement of the floor of the pharynx. Crocodiles close their nostrils when submerged, so olfaction underwater is unlikely. Underwater food detection is presumably gustatory and tactile.Schwenk, K. (2008). Comparative anatomy and physiology of chemical senses in nonavian aquatic reptiles. In, Sensory Evolution on the Threshold: Adaptations in Secondarily Aquatic Vertebrates. J.G.M Hans Thewissen and S. Nummels (Eds). University of California Press, Berkeley. pp. 65–81
Cranial: The upper and lower jaws are covered with sensory pits, visible as small, black speckles on the skin, the crocodilian version of the lateral line organs seen in fish and many amphibians, though arising from a completely different origin. These pigmented nodules encase bundles of Axon innervated beneath by branches of the trigeminal nerve. They respond to the slightest disturbance in surface water, detecting vibrations and small pressure changes as small as a single drop. This makes it possible for crocodiles to detect prey, danger and intruders, even in total darkness. These sense organs are known as domed pressure receptors (DPRs).
Post-Cranial: While alligators and caimans have DPRs only on their jaws, crocodiles have similar organs on almost every scale on their bodies. The function of the DPRs on the jaws is clear; to catch prey, but it is still not clear what the function is of the organs on the rest of the body. The receptors flatten when exposed to increased osmotic pressure, such as that experienced when swimming in sea water hyperosmotic to the body fluids. When contact between the integument and the surrounding sea water solution is blocked, crocodiles are found to lose their ability to discriminate salinities. It has been proposed that the flattening of the sensory organ in hyperosmotic sea water is sensed by the animal as "touch", but interpreted as chemical information about its surroundings. This might be why in alligators they are absent on the rest of the body.
Crocodiles have the most acidic stomach of any vertebrate. They can easily digest bones, hooves and horns. The BBC TVBBC channel 1 program Inside The Perfect Predator, Thursday 25 March 2010 reported that a Nile crocodile that has lurked a long time underwater to catch prey builds up a large oxygen debt. When it has caught and eaten that prey, it closes its right aortic arch and uses its left aortic arch to flush blood loaded with carbon dioxide from its muscles directly to its stomach; the resulting excess acidity in its blood supply makes it much easier for the stomach lining to secrete more stomach acid to quickly dissolve bulks of swallowed prey flesh and bone. Many large crocodilians swallow stones (called gastroliths or stomach stones), which may act as ballast to balance their bodies or assist in crushing food, similar to grit ingested by birds. Herodotus claimed that Nile crocodiles had a symbiosis with certain birds, such as the Egyptian plover, which enter the crocodile's mouth and pick feeding on the crocodile's blood; with no evidence of this interaction actually occurring in any crocodile species, it is most likely mythical or allegorical fiction.
A long saltwater crocodile has been confirmed as having the strongest bite ever recorded for an animal in a laboratory setting. It was able to apply a bite force value of , and thus surpassed the previous record of made by a long American alligator. Based on regression of mean body mass versus mean bite force, the bite force of a saltwater crocodile with an estimated mass of was estimated at between and . The study, led by Dr. Gregory M. Erickson, also shed light on the larger, extinct species of . Since crocodile anatomy has barely changed over the last 200 million years, current data on modern crocodilians can be used to estimate the bite force of extinct species. An Deinosuchus would apply a force of , nearly twice that of the latest, higher bite force estimations of Tyrannosaurus ().Crispian Scully, (2002) Oxford Handbook of Applied Dental Sciences, Oxford University Press – P156 The extraordinary bite of crocodilians is a result of their anatomy. The space for the jaw muscle in the skull is very large, which is easily visible from the outside as a bulge at each side. The muscle is so stiff, it is almost as hard as bone to touch, as if it were the continuum of the skull. Another trait is that most of the muscle in a crocodile's jaw is arranged for clamping down. Despite the strong muscles to close the jaw, crocodiles have extremely small and weak muscles to open the jaw. Crocodiles can thus be subdued for study or transport by duct tape their jaws or holding their jaws shut with large cut from automobile tire.
Crocodiles may possess a homing instinct. In northern Australia, three rogue saltwater crocodiles were relocated by helicopter, but returned to their original locations within three weeks, based on data obtained from tracking devices attached to them.
In captivity, some individuals are claimed to have lived for over a century. A male crocodile lived to an estimated age of 110–115 years in a Russian zoo in Yekaterinburg. Named Kolya, he joined the zoo around 1913 to 1915, fully grown, after touring in an animal show, and lived until 1995. A male freshwater crocodile lived to an estimated age of 120–140 years at the Australia Zoo. Known affectionately as "Mr. Freshie", he was rescued around 1970 by Bob Irwin and Steve Irwin, after being shot twice by hunters and losing an eye as a result, and lived until 2010. Crocworld Conservation Centre, in Scottburgh, South Africa, claims to have a male Nile crocodile that was born in 1900. Named Henry, the crocodile is said to have lived in Botswana along the Okavango River, according to centre director Martin Rodrigues.
Crocodiles are also the most vocal of all reptiles, producing a wide variety of sounds during various situations and conditions, depending on species, age, size and sex. Depending on the context, some species can communicate over 20 different messages through vocalizations alone. Some of these vocalizations are made during social communication, especially during territorial displays towards the same sex and courtship with the opposite sex; the common concern being reproduction. Therefore most conspecific vocalization is made during the breeding season, with the exception being year-round territorial behaviour in some species and quarrels during feeding. Crocodiles also produce different distress calls and in aggressive displays to their own kind and other animals; notably other predators during interspecific predatory confrontations over carcasses and terrestrial kills.
Specific vocalisations include —
periods range from a few weeks up to six months. A hole nest is usually excavated in sand and a mound nest is usually constructed out of vegetation. Females can build or dig several trial nests which appear incomplete and abandoned later. Females are highly protective of their nests and young. Crocodile do not have sex chromosomes, and unlike humans, sex is not determined genetically. Sex is determined by temperature, where at or less most hatchlings are females and at , offspring are of both sexes. A temperature of gives mostly males whereas above in some species continues to give males, but in other species resulting in females, which are sometimes called high-temperature females. Temperature also affects growth and survival rate of the young, which may explain the sexual dimorphism in crocodiles. The average incubation period is around 80 days, and also is dependent on temperature and species that usually ranges from 65 to 95 days. The eggshell structure is very conservative through evolution but there are enough changes to tell different species apart by their eggshell microstructure.
At the time of hatching, the young start calling within the eggs. They have an egg-tooth at the tip of their snouts, which is developed from the skin, and that helps them pierce out of the shell. Hearing the calls, the female usually excavates the nest and sometimes takes the unhatched eggs in her mouth, slowly rolling the eggs to help the process. The young is usually carried to the water in the mouth. She would then introduce her hatchlings to the water and even feed them. 8 November 2008 The mother would then take care of her young for over a year before the next mating season. In the absence of the mother crocodile, the father would act in her place to take care of the young. 3 February 2012 However, even with a sophisticated Paternal care, young crocodiles have a very high mortality rate due to their vulnerability to predation. A group of is called a pod or crèche and may be protected for months.
Vladimir Dinets of the University of Tennessee, observing crocodile's use of twigs as bait was inconclusive.
Crocodile meat is consumed in some countries, such as Australia, Ethiopia, Thailand, South Africa, China, and Cuba (in pickled form). It is also occasionally eaten as an "exotic" delicacy in the western world. Cuts of meat include backstrap and tail fillet.
Due to high demand for crocodile products, TRAFFIC states that 1,418,487 Nile Crocodile skins were exported from Africa between 2006 and 2015.
Crocodiles appear in different forms in Hinduism. Varuna, a Vedic and Hindu god, rides a part-crocodile makara; his consort Varuni rides a crocodile. Similarly the goddess personifications of the Ganges and Yamuna rivers are often depicted as riding crocodiles. Also in India, in Goa, crocodile worship is practised, including the annual Mannge Thapnee ceremony.
Sikh warriors known as nihang also have connections with crocodiles. Nihang may come from the Persian word for a mythical sea creature (). The term owes its origin to Mughal Empire historians, who compared the ferocity of the Akali with that of crocodiles. In Sikhism however, Akali refers to the immortal army of Akal (god).
In Latin America, Cipactli was the giant earth crocodile of the Aztec and other Nahua peoples.
The name of Surabaya, Indonesia, is locally believed to be derived from the words " suro" (shark) and " boyo" (crocodile), two creatures which, in a local mythology, fought each other in order to gain the title of "the strongest and most powerful animal" in the area. It was said that the two powerful animals agreed for a truce and set boundaries; that the shark's domain would be in the sea while the crocodile's domain would be on the land. However one day the shark swam into the river estuary to hunt; this angered the crocodile, who declared it his territory. The Shark argued that the river was a water-realm which meant that it was shark territory, while the crocodile argued that the river flowed deep inland, so it was therefore crocodile territory. The two animals bit each other and a ferocious fight ensued. Finally the shark, badly bitten, fled to the open sea, and in the end the crocodile ruled the estuarine area that today is the city. Another source alludes to a Jayabaya prophecy—a 12th-century psychic king of Kediri Kingdom—as he foresaw a fight between a giant white shark and a giant white crocodile taking place in the area. This is sometimes interpreted as a foretelling of the Mongol invasion of Java, a major conflict between the forces of the Kublai Khan, Mongol Empire ruler of China, and those of Raden Wijaya's Majapahit in 1293. The two animals are now used as the city's symbol, with the two facing and circling each other, as depicted in a statue appropriately located near the entrance to the Surabaya Zoo (see photo on the Surabaya page).
In the UK, a row of schoolchildren walking in pairs, or two by two is known as "crocodile".Martin H. Manser, Turton and Nigel D. Turton Angela Brazil
In politics, a crocodile has been used in different countries to describe Politician and political groups, sometimes as a compliment for a leader seen as strong and strategic, while in others it is a negative label linked to corruption, greed, or Predation.
Taxonomy and phylogeny
Crocodylidae was named as a family by Georges Cuvier in 1807. It belongs to the larger superfamily Crocodyloidea, which also includes additional extinct crocodile relatives. These all belong to the order Crocodilia, which also includes alligators and gharials.
Phylogeny
Recent molecular studies using DNA sequencing have shown crocodiles to be more closely related to the rather than to , contrary to prior theories based on morphological studies alone.
Species
A total of 18 extant taxon species have been recognized. Further genetic study is needed for the confirmation of proposed species under the genus Osteolaemus.
American crocodile () Throughout the Caribbean Basin, including many of the and South Florida. A larger sized species, with a greyish colour and a prominent V-shaped snout. Prefers brackish water, but also inhabits lower stretches of and true marine biology environments. This is one of the rare species that exhibits regular sea-going behaviour, which explains the great distribution throughout the Caribbean. It is also found in such as Lago Enriquillo, in the Dominican Republic, which has one of the largest populations of this species. Diet consists mostly of aquatic and terrestrial vertebrates. Classified as Vulnerable, but certain local populations under greater threat. Crocodylus halli ( Crocodylus halli) The island of New Guinea, south of the New Guinea Highlands A smaller species that closely resembles and was long classified under the New Guinea crocodile, which it is now considered to be genetically distinct from. It lives south of the mountain barrier that divides the two species' ranges. It can be physically distinguished from the New Guinea crocodile by its shorter maxilla and enlarged elements. Cranial elements can still widely vary within the species, with populations from Lake Murray having much wider heads than those from the Aramia River. Orinoco crocodile () Colombia and Venezuela This is a large species with a relatively elongated snout and a pale tan coloration with scattered dark brown markings. Lives primarily in the Orinoco. Despite having a rather narrow snout, preys on a wide variety of vertebrates, including large mammals. It is a Critically Endangered species. Freshwater crocodile () Northern Australia A smaller species with a narrow and elongated snout. It has light brown coloration with darker bands on body and tail. Lives in with considerable distance from the sea, to avoid confrontations with saltwater crocodiles. Feeds mostly on fish and other small vertebrates. Philippine crocodile () Endemic to the Philippines This is a relatively small species with a rather broader snout. It has heavy dorsal armour and a golden-brown colour that darkens as the animal matures. Prefers freshwater habitats and feeds on a variety of small to medium sized vertebrates. This species is Critically Endangered and the most severely threatened species of crocodile.(2011-09-06). "Pictures: Biggest Crocodile Ever Caught?". National Geographic Daily News. Morelet's crocodile () Atlantic regions of Mexico, Belize and Guatemala A small to medium sized crocodile with a rather broad snout. It has a dark greyish-brown colour and is found in mostly various freshwater habitats. Feeds on mammals, birds and reptiles. It is listed as Least Concern. Nile crocodile () Sub-saharan Africa A large and aggressive species with a broad snout, especially in older animals. It has a dark bronze coloration and darkens as the animal matures. Lives in a variety of freshwater habitats but is also found in brackish water. It is an apex predator that is capable of taking a wide array of vertebrates, including large ungulates and other predators. This species is listed as Least Concern. New Guinea crocodile ( The island of New Guinea, north of the New Guinea Highlands A smaller species of crocodile with a grey-brown colour and dark brown to black markings on the tail. The young have a narrower V-shaped snout that becomes wider as the animal matures. Prefers freshwater habitats, even though is tolerant to salt water, in order to avoid competition and predation by the saltwater crocodile. This species feeds on small to mid-sized vertebrates. Mugger crocodile () The Indian subcontinent and surrounding countries This is a modest sized crocodile with a very broad snout and an alligator-like appearance. It has dark-grey to brown coloration. Enlarged scutes around the neck make it a heavily armoured species. Prefers slow moving , and . It can also be found in coastal swamps but avoids areas populated by saltwater crocodiles.[3] . Crocodile Species List. Retrieved on 13 April 2013. Feeds on a wide array of vertebrates. Saltwater crocodile () Throughout , Northern Australia and surrounding waters The largest on average living reptile and most aggressive of all crocodiles. It is a big-headed species and has a relatively broad snout, especially when older. The coloration is pale yellow with black stripes when young but dark greenish-drab coloured as adults. Lives in brackish and marine biology environments as well as lower stretches of . This species has the greatest distribution of all crocodiles. Tagged specimens showed long-distance marine travelling behaviour. It is the apex predator throughout its range and preys on virtually any animal within its reach. It is classified as Least Concern but with several populations under greater risk. Borneo crocodile ( Crocodylus raninus) Island of Borneo in Southeast Asia A freshwater species of crocodile that has been considered a synonym of the saltwater crocodile. Cuban crocodile () Found only in the Zapata Swamp and Isle of Youth of Cuba It is a small but extremely aggressive species of crocodile that prefers freshwater swamps. The coloration is vibrant even as adults and the scales have a "pebbled" appearance. It is a relatively terrestrial species with agile locomotion on land, and sometimes displays terrestrial hunting. The snout is broad with a thick upper-jaw and large teeth. The unique characteristics and fossil record indicates a rather specialized diet in the past, preying on megafauna such as the giant sloth. This species sometimes displays pack-hunting behaviour, which might have been the key to hunting large species in the past, despite its small size. Today most prey are small to medium sized vertebrates. It is Critically Endangered, and the remaining wild population is under threat of hybridization. Siamese crocodile () Indonesia, Brunei, East Malaysia and southern Indochina A fairly small crocodile that prefers freshwater habitats. It has a relatively broad snout and olive-green to dark green coloration. It feeds on a variety of small to mid-sized vertebrates. Listed as Critically Endangered, but might be already extinct in the wild; status is unknown. West African crocodile () Western Africa and Central Africa Recent studies revealed that this is distinct species from the larger Nile crocodile. Nile crocodile is two species, Nature.com It has a slightly narrower snout and is much smaller compared to its larger cousin. Osborn's dwarf crocodile () Western Africa It is a heavily armoured species with uniform black coloration in adults, while juveniles have a lighter brown banding. Lives in the tropical forests of Western Africa. Feeds on small vertebrates and large aquatic invertebrates. It is a fairly terrestrial species and exhibits terrestrial hunting, especially at night. Dwarf crocodile () Western Africa It belongs to its own monotypic genus; however, new studies indicate there might be two or even three distinct species. It is a heavily armoured species with uniform black coloration in adults, while juveniles have a lighter brown banding. Lives in the tropical forests of Western Africa. Feeds on small vertebrates and large aquatic invertebrates. It is a fairly terrestrial species and exhibits terrestrial hunting, especially at night. This species is classified as Vulnerable. West African slender-snouted crocodile () Western Africa A medium sized species with a narrow and elongated snout. Lives in freshwater habitats within tropical forests of the continent. Feeds mostly on fish but also other small to medium sized vertebrates. It is a Critically Endangered species. Central African slender-snouted crocodile ( Mecistops leptorhynchus) Central Africa A medium sized species found in watery areas in dense rainforest. Feeds largely on fish. Insufficient conservation data, but was classified as Critically Endangered when lumped with M. cataphractus, although M. leptorhynchus is doing better in its home range.
Characteristics
Apart from the advantage conferred by its sufficiently large size relative to other animals in the ecosystem, other physical traits contribute a crocodile's position as predator. Its external morphology is a sign of its aquatic animal and predatory lifestyle. Its streamlined body enables it to swim swiftly; it also tucks its feet to the side while swimming, making it faster by decreasing water resistance. Crocodiles have webbed feet which, though not used to propel them through the water, allow them to make fast turns and sudden moves in the water or initiate swimming. Webbed feet are an advantage in shallow water, where the animals sometimes move around by walking. Crocodiles have a palatal flap, a rigid tissue at the back of the mouth that blocks the entry of water. The palate has a special path from the nostril to the glottis that bypasses the mouth. The nostrils are closed during submergence.
Size
Size greatly varies among species, from the dwarf crocodile to the saltwater crocodile. Species of the dwarf crocodile Osteolaemus grow to an adult size of just ,[6]. Crocodilian Species List. Retrieved on 14 April 2012 whereas the saltwater crocodile can grow to sizes over and weigh over .Guinness Book of World Records. Retrieved on 8 April 2013. Several other large species can reach over long and weigh over . Crocodilians show pronounced sexual dimorphism, with males growing much larger and more rapidly than females. Despite their large adult sizes, crocodiles start their lives at around long. The largest species of crocodile is the saltwater crocodile, found in eastern India, northern Australia, throughout South-east Asia, and in the surrounding waters.
Teeth
Crocodiles are ; they are able to replace each of their 80 teeth up to 50 times in their 35- to 75-year lifespan. Next to each full-grown tooth, there is a small replacement tooth and an odontogenic stem cell in the dental lamina in standby that can be activated if required. Specialized stem cell niche enables repetitive renewal of alligator teeth
Biology and behaviour
Crocodilians are more closely related to birds and dinosaurs than to most animals classified as reptiles, the three families being included in the group Archosauria ('ruling reptiles'). Despite their prehistoric look, crocodiles are among the more biologically complex reptiles. Unlike other reptiles, a crocodile has a cerebral cortex and a four-chambered heart. Crocodilians also have the functional equivalent of a diaphragm by incorporating muscles used for aquatic locomotion into respiration. are present in the tongues of crocodiles and they have a pore opening on the surface of the tongue, a trait that separates them from alligators. Salt glands are dysfunctional in Alligatoridae. Their function appears to be similar to that of salt glands in . Crocodiles do not have sweat glands and release heat through their mouths. They often sleep with their mouths open and may pant like a dog. Four species of freshwater crocodile climb trees to bask in areas lacking a shoreline.
Senses
Crocodiles have acute senses, an evolutionary advantage that makes them successful predators. The eyes, ears and nostrils are located on top of the head, allowing the crocodile to lie low in the water, almost totally submerged and hidden from prey.
Vision
Crocodiles have very good night vision, and are mostly Nocturnality hunters. They use the disadvantage of most prey animals' poor nocturnal vision to their advantage. The light receptors in crocodilians' eyes include Cone cell and numerous Rod cells, so it is assumed all crocodilians can see colours. Crocodiles have vertical-slit shaped pupils, similar to those of domestic cats. One explanation for the evolution of slit pupils is that they exclude light more effectively than a circular pupil, helping to protect the eyes during daylight. On the rear wall of the eye is a tapetum lucidum, which reflects incoming light back onto the retina, thus utilizing the small amount of light available at night to best advantage. In addition to the protection of the upper and lower eyelids, crocodiles have a nictitating membrane (sometimes called a "third eye-lid") that can be drawn over the eye from the inner corner while the lids are open. The eyeball surface is thus protected under the water while a certain degree of vision is still possible.
Olfaction
Crocodilian Olfaction is also very well developed, aiding them to detect prey or animal carcasses that are either on land or in water, from far away. It is possible that crocodiles use olfaction in the egg prior to hatching.
Hearing
Crocodiles can hear well; their Eardrum are concealed by flat flaps that may be raised or lowered by muscles.
Touch
The touch sensors, concentrated in crocodile skin, can be thicker than those in human fingerprints. Crocodiles can feel the touch on their skin.
Hunting and diet
Crocodiles are , waiting for fish or land animals to come close, then rushing out to attack. Crocodiles mostly eat fish, , , , , , and , and they occasionally cannibalize smaller crocodiles. What a crocodile eats varies greatly with species, size and age. From the mostly fish-eating species, like the slender-snouted and freshwater crocodiles, to the larger species like the Nile crocodile and the saltwater crocodile that prey on large mammals, such as Bovinae, deer and wild boar, diet shows great diversity. Diet is also greatly affected by the size and age of the individual within the same species. All young crocodiles hunt mostly invertebrates and small fish, gradually moving on to larger prey. Being (cold-blooded) predators, they have a very slow metabolism, so they can survive long periods without food. Despite their appearance of being slow, crocodiles have a very fast strike and are top in their environment, and various species have been observed attacking and killing other such as and . Crocodiles are also known to be aggressive who feed upon carrion and steal from other predators. Evidence suggests that crocodiles also feed upon fruits, based on the discovery of seeds in stools and stomachs from many subjects as well as accounts of them feeding. 13 November 2013 27 August 2013
Bite
Since they feed by grabbing and holding onto their prey, they have Evolution sharp teeth for piercing and holding onto flesh, and powerful muscles to close the jaws and hold them shut. The teeth are not well-suited to tearing flesh off of large prey items as are the dentition and claws of many mammalian carnivores, the hooked bills and talons of raptorial birds, or the serrated teeth of sharks. However, this is an advantage rather than a disadvantage to the crocodile since the properties of the teeth allow it to hold onto prey with the least possibility of the prey animal escaping. Cutting teeth, combined with the exceptionally high bite, would pass through flesh easily enough to leave an escape opportunity for prey. The jaws can bite down with immense force, by far the strongest bite of any animal. The force of a large crocodile's bite is more than , which was measured in a Nile crocodile, in the field;National Geographic documentary; "Bite Force", Brady Barr. comparing to for a Rottweiler, for a hyena, for an American alligator, and for the largest confirmed great white shark.
Locomotion
Crocodiles can move quickly over short distances, even out of water. The land speed record for a crocodile is measured in a galloping Australian freshwater crocodile. Maximum speed varies between species. Some species can gallop, including Cuban crocodiles, Johnston's crocodiles, New Guinea crocodiles, Dwarf crocodile, and even small Nile crocodiles. The fastest means by which most species can move is a "belly run", in which the body moves in a snake-like (sinusoidal) fashion, limbs splayed out to either side paddling away frantically while the tail whips to and fro. Crocodiles can reach speeds of when they "belly run", and often faster if slipping down muddy riverbanks. When a crocodile walks quickly, it holds its legs in a straighter and more upright position under its body, which is called the "high walk". This walk allows a speed of up to 5 km/h.9781842739648, Marks and Spencer p.l.c. ISBN 9781842739648
Longevity
Measuring crocodile age is unreliable, although several techniques are used to derive a reasonable guess. The most common method is to measure lamellar growth rings in bones and teeth—each ring corresponds to a change in growth rate which typically occurs once a year between dry and wet seasons. Bearing these inaccuracies in mind, it can be safely said that all crocodile species have an average lifespan of at least 30–40 years, and in the case of larger species an average of 60–70 years. The oldest crocodiles appear to be the largest species. C. porosus is estimated to live around 70 years on average, with limited evidence of some individuals exceeding 100 years.
Social behaviour and vocalization
Crocodiles are the most social of reptiles. Even though they do not form social groups, many species congregate in certain sections of , tolerating each other at times of feeding and Body temperature. Most species are not highly territorial, with the exception of the saltwater crocodile, which is a highly territorial and aggressive species: a mature, male saltwater crocodile will not tolerate any other males at any time of the year, but most other species are more flexible. There is a certain form of hierarchy in crocodiles: the largest and heaviest males are at the top, having access to the best basking site, while females are priority during a group feeding of a big kill or carcass. A good example of the hierarchy in crocodiles would be the case of the Nile crocodile. This species clearly displays all of these behaviours. Studies in this area are not thorough, however, and many species are yet to be studied in greater detail. are also known to show toleration in group feedings and tend to congregate in certain areas. However, males of all species are aggressive towards each other during mating season, to gain access to females.
(1991). 9780679735663, Vintage Books. ISBN 9780679735663
Reproduction
Mating
Courtship takes place in a series of behavioural interactions that include a variety of snout rubbing and submissive display that can take a long time. Mating always takes place in water, where the pair can be observed mating several times.
Egg-laying & nesting
Egg-laying usually takes place at night and about 30–40 minutes, which are laid in either holes or mound , depending on species. The eggs are hard shelled, but translucent at the time of egg-laying. Depending on the species of crocodile, 7 to 95 eggs are laid. may play a role in calcium storage for eggshell formation.
Cognition
Crocodiles possess some advanced cognitive abilities. 6 December 2013 9 December 2013 Crocodiles cooperatively hunt. Large numbers of crocodiles swim in circles to trap fish and take turns snatching them. In hunting larger prey, crocodiles swarm in, with one holding the prey down as the others rip it apart.
Relationship with humans
Danger to humans
The larger species of crocodiles are very dangerous to humans, mainly because of their ability to strike before the person can react. The saltwater crocodile and Nile crocodile are the most dangerous, killing hundreds of people each year in parts of Southeast Asia and Africa. The mugger crocodile and American crocodile are also dangerous to humans.
Crocodile products
Crocodiles are protected in many parts of the world, but are also farmed commercially. Their hides are tanned and used to make leather goods such as shoes and ; crocodile meat is also considered a delicacy. The most commonly farmed species are the saltwater and Nile crocodiles, while a hybrid of the saltwater and the rare Siamese crocodile is also bred in Asian farms. Farming has resulted in an increase in the saltwater crocodile population in Australia, as eggs are usually harvested from the wild, so landowners have an incentive to conserve their habitat. Crocodile leather can be made into goods such as wallets, briefcases, purses, handbags, belts, hats, and shoes. Crocodile oil has been used for various purposes. Crocodiles were eaten by Vietnamese while they were taboo and off limits for Chinese. Vietnamese women who married Chinese men adopted the Chinese taboo.
Crocodile hunting and conservation
Aboriginal Australians harvested eggs and hunted crocodiles in a sustainable way for many thousands of years. The Brinkin people (aka Marrithiyal) of the Daly River in the Northern Territory (NT) used and bamboo, and even their own hands to capture crocodiles for food. After settlement of northern Australia, in the late-19th and early 20th centuries, non-Indigenous people killed individual crocodiles, mostly by locals to protect the population, or novelty-seeking visitors, or just opportunistically, so numbers were not noticeably reduced. From the 1930s, commercial hunting began, with Aboriginal people often employed to kill the crocodiles using traditional methods. From the 1940s to the 1960s, hunting began on a larger scale using .303 rifles. They were hunted for leather, with the skins shipped to plants in capital cities. Western Australia banned hunting freshwater crocodiles in 1962 and saltwater crocodiles in 1970, while NT bans were brought in 1964 and 1971; Queensland did not pass such legislation. The federal government later banned the export of crocodile skins, which brought commercial hunting to an end in Queensland. They have been a protected species since the 1970s, when numbers were down to approximately 3,000 in the NT at the lowest estimate. In 2021, after several attacks on humans by the "salties" and an estimated population of around 200,000 had been reached, Queensland politician Bob Katter called for the reintroduction of hunting.
In religion and mythology
Crocodiles have appeared in various forms in religions across the world. Ancient Egypt had Sobek, the crocodile-headed god, with his cult-city Crocodilopolis, as well as Taweret, the goddess of childbirth and fertility, with the back and tail of a crocodile. The Jukun shrine in the Wukari Federation, Nigeria is dedicated to crocodiles in thanks for their aid during migration. 24 June 2009 In Madagascar various peoples such as the Sakalava people and Antandroy see crocodiles as ancestor spirits and under local fady often offer them food;Campbell, Gwyn (2012). David Griffiths and the Missionary "History of Madagascar". Leiden, The Netherlands: Brill. .Marty Crump, Livros no Google Play
Eye of Newt and Toe of Frog, Adder's Fork and Lizard's Leg: The Lore and Mythology of Amphibians and Reptiles, University of Chicago Press, 16 November 2015 in the case of the latter at least a crocodile features prominently as an ancestor deity.An Athenæum article makes a claim of a specific crocodile deity named textually as "Jacaret", but less biased sources do not mention this theonym in any capacity.
In language and as symbols
The term "crocodile tears" (and equivalents in other languages) refers to a false, insincere display of emotion, such as a hypocrite crying fake tears of grief. It is derived from an ancient anecdote that crocodiles weep in order to lure their prey, or that they Crying for the victims they are eating, first told in the Bibliotheca by Photios I of Constantinople. The story is repeated in bestiaries such as De bestiis et aliis rebus. This tale was first spread widely in English in the stories of the Travels of Sir John Mandeville in the 14th century, and appears in several of Shakespeare's plays. In fact, crocodiles can and do generate tears, but they do not actually cry. Britton, Adam (n.d.). Do crocodiles cry 'crocodile tears'? Crocodilian Biology Database. Retrieved 13 March 2006 from the Crocodile Specialist Group, Crocodile Species List, FAQ.
Fashion logos
The French clothing company Lacoste features a crocodile in its logo. The American shoe company Crocs also uses this imagery in its logo.
See also
Notes
Further reading
External links
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